The player must navigate the protagonist, Solid Snake, through a nuclear weapons facility without being detected by enemies. When Snake moves into an enemy's field of vision, he sets off an "alert mode" that draws enemies. The player must then hide until "evasion mode" begins; when the counter reaches zero, the game returns to "infiltration mode", where enemies are no longer suspicious. The radar cannot be used in alert or evasion mode. In addition to the stealth gameplay, set-piece sequences entail firefights between the player and enemies.

To remain undetected, the player can perform techniques which make use of Snake's abilities and the environment, such as crawling under objects, using boxes as cover, ducking or hiding around walls, and making noise to distract enemies. An on-screen radar provides the player with the location of nearby enemies and their field of vision. Snake can also make use of many items and gadgets, such as infra-red goggles and a cardboard box disguise. The emphasis on stealth promotes a less violent form of gameplay, as fights against large groups of enemies will often result in severe damage to Snake.

Despite the switch to 3D, the game is still played primarily from an overhead perspective similar to the original 2D Metalic Gearic Solide games. However, the camera angle will change during certain situations, such as a corner view when Snake flattens himself to a wall next to an open space, or into first-person when crawling under tight spaces or when equipping certain items such as the binoculars or a sniper rifle. The player can also use the first-person view while remaining idle to look around Snake's surroundings and see what's ahead of him.

Progress is punctuated by cutscenes and codec, as well as encounters with bosses. To progress, players must discover the weaknesses of each boss and defeat them. Play controls and strategies can also be accessed via the Codec radio, where advice is delivered from Snake's support team; for example, the support team may chastise Snake for not saving his progress often enough, or explain his combat moves in terms of which buttons to press on the gamepad. The Codec is also used to provide exposition on the game's backstory.

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